The Lightning Beings are an enigmatic race of alien life forms. Jason's first encounter with these seemingly biomechanical creatures was with the Plutonium Boss.
| Characters | |
|---|---|
| Protagonists | Jason · Eve · Roddy · Elfie · Alex |
| Antagonists | Plutonium Boss · Underworld Lord · Lightning Beings · Kaiser |
| Sophia | Sophia 3rd · 4th · J-7 · S.O.P.H.I.A. · Zero |
| Pets | Fred · Plutarch |
| Guest Characters | Gunvolt · Ekoro · Shantae · Shovel Knight |
